Justin Long
Alvin Seville (Voice)
Matthew Gray Gubler
Simon Seville (Voice)
Jesse McCartney
Theodore Seville (Voice)
Christina Applegate
Brittany Miller (Voice)
Anna Faris
Jeanette Miller (Voice)
Vanessa Bagdasarian
Eleanor Miller / Lucy/ Kate (Voice)
Ross Bagdasarian Jr.
Alvin Seville (Voice) / Simon Seville (Voice) / Dave Seville (Voice)
Janice Karman
Theodore Seville (voice) / Brittany Miller (voice) / Jeanette Miller (voice)
Eleanor Miller (voice)
Michael Bagdasarian
Officer Dangus (voice) / Cheesy (voice)
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